Horses and other equids are odd-toed ungulates of the order Perissodactyla, a relatively ancient group of browsing and grazing animals that first arose less than 10 million years after the dinosaurs became extinct. In the past, this order contained twelve families, but only three families— Equidae (the horse and related species), the tapir and the rhinoceros—have survived to the present day. The earliest equids known as Hyracotherium developed approximately 54 million years ago, during the Eocene period. One of the first true horse species, it had 4 toes on each front foot, and 3 toes on each back foot. the extra toe on the front feet soon disappeared, and by the Pleistocene era, as the horse adapted to a drier, prairie environment, the 2nd and 4th toes disappeared on all feet, and horses became bigger. These side toes first shrunk in size until they have vanished in modern horses.